Prehistory and Anthropology

Grotta dei Santi - Monte Argentario (GR) (Paleolitico medio)

The activities of the Research Unit of Prehistory and Anthropology (RUPA) focus on the study and the reconstruction of prehistoric environments and societies, along a time span from the Paleolithic to the Metal Ages.

The objects of investigation are paleoecology, subsistence strategies, the ability to exploit natural resources, technical knowledge, artefacts, features (e.g., hearths), and evidence of symbolic thinking of prehistoric peoples (ornaments, burials, art, etc.). Hence there many disciplines contributing to the study of prehistory, e.g., paleoanthropology, palethnology, geomorphology, sedimentology, pedology, volcanology, paleontology, zooarchaeology, archaeobotany, paleochronology, paleogenetics, and paleonutrition. Some of them are covered within the DSFTA, others are addressed in collaboration with external research institutes.


The first phase of research involves surveys and excavations in caves, shelters and open-air prehistoric sites that preserve evidence of human occupations and allow investigation of climatic/environmental contexts and of interactions between humans and their coeval environments. Fieldwork has also educational value for both undergraduates and postgraduates.


Laboratory research involves the study of findings and elaboration of data from fieldwork activities.  Artefacts (stone, bone, pottery, etc.) are analysed from a perspective which is technological (raw material supply and characteristics, production techniques, etc.), typological (final form of the objects and their attribution to a cultural phase) and functional (use-wear traces analyzed by SEM and optical and digital microscopy).

The study of faunal remains from hunting, breeding and slaughtering activities carried out by prehistoric people, provides pivotal information on paleoenvironmental patterns and paleoeconomical strategies.


The RUPA is equipped with laboratories for post-excavation activities. There are reference osteological collections for both zooarchaeology and paleoanthropology, a lithoteque and a reference collection for the study of lithic artefacts, and a Hirox KH-7700 3D digital microscope for both the analysis of artefacts and osteological remains.

Lines of research
Ginosa (TA) - Riparo l'Oscurusciuto (Paleolitico Medio)

Our research focuses on the reconstruction of the cultural evolution of Neanderthal groups based on stratigraphic excavations carried out since the ‘60’s in deposits of the Tyrrhenian (Cilento and Tuscany) and Adriatic (Puglia) areas, among the most important in the Mediterranean. Special attention is paid to the dynamics connected to the replacement of the last Neanderthal populations by modern humans (Homo sapiens) at the time of their arrival and spread in Europe.


Mariotti Lippi M., Aranguren B., Arrighi A., Attolini D., Benazzi S., Boschin F., Florindi S., Moroni A., Negrino F., Pallecchi P., Pisaneschi L., Riel-Salvatore J., Ronchitelli A., Revedin A. 2023 New evidence of plant food processing in Italy before 40ka. Quaternary Science Reviews 312, 108161.


Rossini M., Marciani G., Arrighi S., Pereira Santos M.C., Spagnolo V., Ronchitelli A., Benazzi S., Moroni A. 2022. Less is more! Uluzzian technical behaviour at the cave site of Castelcivita (southern Italy). Journal of Archaeological Science: Reports 44, 103494.


Marciani, G., Spagnolo, V., Martini I., Casagli A., Sulpizio R., Aureli, D., Boscato, P., Ronchitelli A., Boschin, F. 2020. Neanderthal occupation during the tephra fall-out: Technical and hunting behaviours, sedimentology and settlement patterns in SU 14 of Oscurusciuto rock shelter (Ginosa, southern Italy). Archaeological and Anthropological Sciences 12: 152.


Spagnolo, V., Crezzini, J., Marciani, G., Capecchi, G., Arrighi, S., Aureli, D., Scaramucci, S., Boschin, F., Moroni, A. 2020. Neandertal camps and hyena dens. Living floor 150A at Grotta dei Santi (Monte Argentario, Tuscany, Italy). Journal of Archaeological Science: Report, 30, 102249


Spagnolo, V., Aureli, D., Martini, I., Ekberg, I., Boschin, F., Crezzini, J., Poggi G., Boscato P., Ronchitelli A. 2020. Short and close in time: overlapped occupation from the layer 56 of the Molare Rock shelter (Southern Italy). Archaeological and Anthropological Sciences, 12, 92.


Spagnolo V., Marciani G., Aureli D., Martini I., Boscato P., Boschin F., Ronchitelli A. 2020. Climbing the time to see Neanderthal behaviour’s continuity and discontinuity: SU 11 of the Oscurusciuto Rockshelter (Ginosa, Southern Italy). Archaeological and Anthropological Sciences 12, 54.


Arrighi S., Moroni A., Tassoni L., Boschin F., Badino F., Bortolini E., Boscato P., Crezzini J., Figus C., Forte M., Lugli F., Marciani G., Oxilia G., Negrino F., Riel-Salvatore J., Romandini M., Peresani M., Spinapolice E.E., Ronchitelli A., Benazzi S. 2020. Bone tools, ornaments and other unusual objects during the Middle to Upper Palaeolithic transition in Italy. Quaternary International 551, 169-187.


Marciani G., Ronchitelli A., Arrighi S., Badino F., Bortolini E., Boscato P., Boschin F., Crezzini J., Delpiano D., Falcucci A., Figus C., Lugli F., Oxilia G., Romandini M., Riel-Salvatore J., Negrino F., Peresani M., Spinapolice E.E., Moroni A., Benazzi S. 2020. Lithic techno-complexes in Italy from 50 to 39 thousand years BP: An overview of lithic technological changes across the Middle-Upper Palaeolithic boundary. Quaternary International 551, 123-149.


Fiore I., Gala M., Boschin F., Crezzini J., Tagliacozzo A., Moroni A., 2020. Archeozoology and taphonomy of bird remains from Grotta di Castelcivita (Salerno, Italy) and clues for human-bird interactions. Quaternary International 551, 224-242.


Sano K., Arrighi S., Stani C., Aureli D., Boschin F., Fiore I., Spagnolo V., Ricci S., Crezzini J., Boscato P., Gala M., Tagliacozzo A., Birarda G., Vaccari L., Ronchitelli A., Moroni A., Benazzi S. 2019. The earliest evidence for mechanically delivered projectile weapons in Europe. Nature Ecology and Evolution 3, 1409-1414.


Moroni A., Boschian G., Crezzini J, Montanari-Canini G, Marciani G, Capecchi G, Arrighi S, Aureli S, Aureli D, Berto C, Freguglia M, Araujo A, Scaramucci S, Hublin JJ, Lauer T, Benazzi S, Parenti F, Bonato M, Ricci S, Talamo S, Segre AG, Boschin F, Spagnolo V. 2019. Late Neandertals in central Italy. High-resolution chronicles from Grotta dei Santi (Monte Argentario - Tuscany). Quaternary Science Reviews 217, 130-151.


Moroni A., Ronchitelli A., Arrighi S., Aureli D., Bailey S., Boscato P., Boschin F., Capecchi G., Crezzini J., Douka K., Marciani G., Panetta D., Ranaldo F., Ricci S., Scaramucci S., Spagnolo V., Benazzi S., Gambassini P. 2018. Grotta del Cavallo (Apulia-Southern Italy). The Uluzzian in the mirror. Journal of anthropological sciences 96.


Higham T., Douka K., Wood R., Ramsey C.B., Brock F. et al. 2014 The timing and spatiotemporal patterning of Neanderthal disappearance. Nature 512, 306-309.


Benazzi S., Douka K., Fornai C., Bauer C.C., Kullmer O., Svoboda J., Pap I., Mallegni F., Bayle P., Coquerelle M., Condemi S., Ronchitelli A., Harvati K., Weber G.W. 2011. Early dispersal of modern humans in Europe and implications for Neanderthal behaviour. Nature 479, 525-528.

Grotta Paglicci (Rignano Garganico-FG): arte e pitture parietali paleolitiche

This study mainly focuses on Palaeolithic hunter-gatherers who lived in the Italian Peninsula during the last climatic cold oscillations. The RUPA team has been investigating for decades important stratigraphic series located in central-southern Italy and is collaborating with other institutions for evidence from northern regions. The main topics of this research line are:

I.            “Italy as a refugium during the Last Glacial Maximum”;

II.           “Ecological and cultural adaptation of Palaeolithic hunter-gatherers to the last harsh climatic oscillations”;

III.          “Mobility and migratory patterns of Upper Palaeolithic populations”;

IV.          “First steps of mammal domestication ”.


Posth C., Yu H., Ghalichi A., Rougier H., Crevecoeur C., et al. 2023 Palaeogenomics of Upper Palaeolithic to Neolithic European hunter-gatherers. Nature 615, 117-126.


Quagliariello A., Modi A., Innocenti G., Zaro V., Conati Barbaro C., Ronchitelli A., Boschin F., Cavazzuti C., Dellù E., Radina F., Sperduti A., Bondioli L., Ricci S., Lognoli M., Belcastro M.G., Mariotti V., Caramelli D., Mariotti Lippi M., Cristiani E., Martino M.E., Muntoni I.M., Lari M. 2022. Ancient oral microbiomes support gradual Neolithic dietary shifts towards agriculture. Nature Communications 13, 6927. 10.1038/s41467-022-34416-0


Boschin F., Bernardini F., Pilli E., Vai S., Zanolli C., Tagliacozzo A., Fico R., Fedi M., Corny J., Dreossi D., Lari M., Modi A., Vergata C., Tuniz C., Moroni A., Boscato P., Caramelli D., Ronchitelli A. 2020. The first evidence for Late Pleistocene dogs in Italy. Scientific Reports 10, 13313.


Duches R., Nannini N., Fontana A., Boschin F., Crezzini J., Peresani M. 2020. Experimental and archaeological data for the identification of projectile impact marks on small-sized mammals. Scientific reports 10, 9092. 10.1038/s41598-020-66044-3


Lugli F., Cipriani A., Capecchi G., Ricci S., Boschin F., Boscato P., Iacumin P., Badino F., Mannino M.A., Talamo S., Richards M.P., Benazzi S., Ronchitelli A. 2019. Strontium and stable isotope evidence of human mobility strategies across the Last Glacial Maximum in southern Italy. Nature Ecology and Evolution 3 (6), 905-911.


Martini I., Ronchitelli A., Arrighi S., Capecchi G., Ricci S., Scaramucci S., Spagnolo V., Gambassini P., Moroni A. 2018. Cave clastic sediments as a tool for refining the study of human occupation of prehistoric sites: insights from the cave site of La Cala (Cilento, southern Italy). Journal of Quaternary Science 33, 586-596.


Fu Q., Posth, C., Hajdinjak M., Petr M., Swapan M., et al. 2016 The genetic history of Ice Age Europe. Nature 534, 200-205. 10.1038/nature17993.


Posth C., Renaud G., Mittnik A., Drucker D., Rougier H., et al. 2016 Pleistocene mitochondrial genomes suggest a single major dispersal of non-africans and a late glacial population turnover in Europe. Current Biology 26, 827-833.


Lippi M.M., Foggi B., Aranguren B., Ronchitelli A., Revedin A. 2015. Multistep food plant processing at Grotta Paglicci (Southern Italy) around 32,600 cal B.P. PNAS 112, 12075-12080.

Gorgo del Ciliegio (Sansepolcro-AR) media Età del Bronzo

This research is aimed at the reconstruction of ancient landscapes and socioeconomic patterns developing in the Upper Tiber Valley from the Middle Palaeolithic to the Iron age.


Moroni A., Spagnolo V., Crezzini J., Boschin F., Benvenuti M., Gardin S., Cipriani S., Arrighi S. 2020. Settlement, space organization and land-use of a small Middle Bronze Age community of central Italy. The case study of Gorgo del Ciliegio (Arezzo-Tuscany). Quaternary International 539, 122-137.


Arrighi, S., Moroni, A., 2016. Socio-economic relations and settlement dynamics in the Upper Tiber valley during the Bronze Age, Rivista di Scienze Preistoriche, LXVI, 181-193.

Moroni, A., Abati, F., Baldanza, A., Coltorti, M., De Angelis, M.C., Mancini, S., Moroni, B., Pieruccini, P., 2011. L'alto e medio bacino del Tevere durante il Paleolitico medio. Ipotesi sul popolamento e la mobilità dei gruppi di cacciatori-raccoglitori neandertaliani. Bollettino di Archeologia on line, Direzione Generale per le Antichità II 2-3, 171-201.


Moroni A. (a cura di), 2013. Caprese prima di Michelangelo. Edizioni Kappa, 96 p.


Moroni Lanfredini A. e Laurenzi G.P. (a cura di), 2011. Pietralba – Indagine multidisciplinare su alcuni manufatti rupestri dell’Alta Valtiberina. Aboca Edizioni, 139 p.


Moroni, A., Benvenuti, M., 2010. L'Alta Valtiberina toscana. Preistoria e Protostoria di un territorio. Le ricerche, gli aspetti culturali, il paleoambiente. IpoTesi di Preistoria 3, 1-26


Iaia C. e Moroni Lanfredini A. (a cura di), 2009. L’Età del Ferro a Sansepolcro – Attività produttive e ambiente nel sito del Trebbio. Aboca Edizioni, 136 p.

La Pietra (Roccastrada-GR): cava di diaspro dell'Età del Rame

This line of research concerns the exploitation of resources and reconstruction of trade networks related to the production of lithic armatures in central-southern Tuscany, with particular focus on the archaeological geo-site of La Pietra in the Farma Valley.


Scaramucci, S., Volpi, V., Costantini, A., Giamello, M., Donati, A., Aranguren, B., Moroni, A., 2016. New challenges in the study of lithic raw materials in central Italy at the dawn of metal working societies: La Pietra and other radiolarite quarry-workshops in Tuscany. Journal of Lithic Studies 3, nr. 2.


Moroni, A., Aranguren, B, Casini, A, Costantini, A., Grandinetti, G., Scaramucci, S., Gambassini, P., 2016. The Prehistoric Quarry of La Pietra (Roccastrada, Grosseto, Tuscany). Copper Age Lithic Workshops and the Production of Bifacial Points in Central Italy, in Tomasso, A., Binder, D., Martino, G., Porraz, G., Simon, P., Naudinot, N. (eds), Actes de la journée de la Société préhistorique française de Nice, 28-29 mars 2013 “Ressources lithiques, productions et transferts entre Alpes et Méditerranée”, Société préhistorique française, (Séances de la Société préhistorique française, 5), Paris, 305-323.


Gambassini P., Marroni G., 1998. Scoperta una cava preistorica di diaspro in Val di Farma. Rassegna di Archeologia 15, 51 – 54.

Laboratorio di Archeozoologia: collezioni di confronto

This line of research is devoted to the investigation of several paleoclimatic and paleoenvironmental proxies (e.g., faunal and botanical remains, sedimentology, paleoclimatic models) and is carried out in collaboration with other national and international institutions. The aim is to reconstruct paleoenvironments and paleoecologycal dynamics in contexts occupied by humans during Prehistory.


Leonardi M., Boschin F., Boscato P., Manica A. 2022. Following the niche: the differential impact of the last glacial maximum on four European ungulates. Communications Biology 5, 1038. 10.1038/s42003-022-03993-7.


Boschin F., Columbu A., Spagnolo V., Crezzini J., Bahain J-J., Falguères C., Benazzi S., Boscato P., Ronchitelli A., Moroni A., Martini I 2022. Human occupation continuity in southern Italy towards the end of the Middle Palaeolithic: a palaeoenvironmental perspective from Apulia. Journal of Quaternary Science 37, 204-216.


Martini I., Baucon A., Boschin F. 2021. Depositional processes and environmental settings in rock shelters: the case of the prehistoric Oscurusciuto site (Southern Italy). Geological Magazine 158, 891-904.


Boschin F., Boscato P., Berto C., Crezzini J., Ronchitelli A. 2018. The palaeoecological meaning of macromammal remains from archaeological sites exemplified by the case study of Grotta Paglicci (Upper Palaeolithic, southern Italy). Quaternary Research 90, 470-482.


Leonardi M., Boschin F., Giampoudakis K., Beyer R.M., Krapp M., Bendrey R., Sommer R., Boscato P., Manica A., Nogues-Bravo D., Orlando L. 2018. Late Quaternary horses in Eurasia in the face of climate and vegetation change. Science advances 4, eaar5589.


Berto C., Boscato P., Boschin F., Luzi E., Ronchitelli A. 2017. Paleoenvironmental and paleoclimatic context during the Upper Palaeolithic (late Upper Pleistocene) in the Italian Peninsula. The small mammal record from Grotta Paglicci. (Rignano Garganico, Foggia, Southern Italy). Quaternary Science Review, 168: 30-41.


Videomicroscopio 3D

The line of research devoted to functional studies includes the traceological study of lithic and bone artefacts (use wear analysis) and the chemical and morphological analysis of residues (residue analysis) to determine how ancient tools were used, while also assessing their state of preservation. Use wear analysis makes use of both the low-power approach (LPA) and the high-power approach (HPA), which are carried out using various optical microscopes (stereomicroscope, metallographic microscope, 3D digital microscope) and the scanning electron microscope (SEM). Residue analysis is performed through both morphological and chemical studies taking advantage of several microscopic and spectroscopic techniques, including 3D digital microscopy, FTIR microscopy, LEXRF microscopy, microbeam PIXE and SEM-EDS. The chemical analysis of ancient residues is carried out in collaboration with several national and international research centres and institutions. The study of the taphonomic alterations of animal hard materials (e.g., shell, bone and antler) exploited for dietary purposes or for the production of artefacts and ornaments is another central aspect of this line of research, involving both the microscopic approach previously described and high-resolution imaging techniques such as micro-computed tomography.


Arrighi S., Bortolini E., Tassoni L., Benocci A., Manganelli G., Spagnolo V., Foresi L.M., Bambini A.M., Lugli F., Badino F., Aureli D., Boschin F., Figus C., Marciani G., Oxilia G., Silvestrini S., Cipriani A., Romandini M., Peresani M., Ronchitelli A., Moroni A., Benazzi S. 2020. Backdating systematic shell ornament making in Europe to 45,000 years ago. Archaeological and Anthropological Sciences 12, 59.

Arrighi S., Moroni A., Tassoni L., Boschin F., Badino F., Bortolini E., Boscato P., Crezzini J., Figus C., Forte M., Lugli F., Marciani G., Oxilia G., Negrino F., Riel-Salvatore J., Romandini M., Peresani M., Spinapolice E.E., Ronchitelli A., Benazzi S. 2020. Bone tools, ornaments and other unusual objects during the Middle to Upper Palaeolithic transition in Italy. Quaternary International 551, 169-187.

Arrighi S., Bazzanella M., Boschin F., Wierer U. 2016. How to make and use a bone “spatula”. An experimental program based on the Mesolithic osseous assemblage of Galgenbühel/Dos de la Forca (Salurn/Salorno, BZ, Italy). Quaternary International, 423: 143-165.

Borgia V., Boschin F., Ronchitelli A., 2016. Bone and antler working at Grotta Paglicci (Rignano Garganico, Foggia, southern Italy). Quaternary International 403, 23-39.

Boschin F., Crezzini J., 2012, Morphometrical Analysis on Cut Marks Using a 3D Digital Microscope. International Journal of Osteoarchaeology, 22: 549-562.

Dominici C., Stani C., Rossini M., Vaccari L., 2022. SR-FTIR microscopy for the study of residues on Palaeolithic stone tools: looking for a methodological protocol. Journal of Physics: Conference Series 2204, 012050.

Moretti E., Arrighi S., Boschin F., Crezzini J., Aureli D., Ronchitelli A., 2015. Using 3D Microscopy to Analyze Experimental Cut Marks on Animal Bones Produced with Different Stone Tools. Ethnobiology Letters 6.

Oxilia G., Peresani M., Romandini M., Matteucci C., Debono Spiteri C., Henry A.G., Schulz D., Archer W., Crezzini J., Boschin F., Boscato P., Jaouen K., Dogandzic T., Broglio A., Moggi-Cecchi J., Fiorenza L., Hublin J-J., Kullmer O., Benazzi S., 2015. Earliest evidence of dental caries manipulation in the Late Upper Palaeolithic. Nature Scientific Reports 5, 12150.

Uomo di Neandertal

Study of human remains of prehistoric, protohistoric, and historical age, carried out in collaboration with other national and international institutions. The research includes anatomical description, taphonomic and paleopathological studies, 3D geometric morphology, tomography and microtomography. Physiognomic reconstructions are also performed.


Tesi C., Ricci S., Levrini L., Giorgetti G., Campagnolo M., Ciliberti R., Fusco R., Larentis O., Licata M. 2023. Defects-related early childhood caries as hints of possible maternal–fetal health issues: Evidence from medieval northern Italy. International Journal of Osteoarchaeology 33, 251-270.


Tesi C., Ricci S., Crezzini J., Badino P., Fusco R., Rossetti C., Gorini I., Licata M. 2022. Wounded to death. Holistic, multimodal reconstruction of the dynamics in a case of multiple perimortem cranial injuries from a medieval site in northern Italy. Journal of Archaeological Science: Reports 46, 103643. 10.1016/j.jasrep.2022.103643.


Posth C., Zaro V., Spyrou M.A., Vai S., Gnecchi-Ruscone G.A., Modi A., Peltzer A., Mötsch A., Nägele K., Vågene Å.J., Nelson E.A., Radzevičiūtė R., Freund C., Bondioli L.M., Cappuccini L., Frenzel H., Pacciani E., Boschin F., Capecchi G., Martini I., Moroni A., Ricci S., Sperduti A., Turchetti M.A., Riga A., Zavattaro M., Zifferero A., Heyne H.O., Fernández-Domínguez E., Kroonen G.J., McCormick M., Haak W., Lari M., Barbujani G., Bondioli L., Bos K.I., Caramelli D., Krause J. 2021. The origin and legacy of the Etruscans through a 2000-year archeogenomic time transect. Science Advances 7, eabi7673.


Sorrentino R., Stephens N.B., Marchi D., DeMars L.J.D. Figus C., Bortolini E., Badino F., Saers J.P.P., Bettuzzi M., Boschin F., Capecchi G., Feletti F., Guarnieri T., May H., Morigi M.P., Parr W., Ricci S., Ronchitelli A., Stock J.T., Carlson K.J., ryan T.M., Belcastro M.G., Benazzi S. 2021. Unique foot posture in Neanderthals reflects their body mass and high mechanical stress. Journal of Human Evolution 161, 103093.


Mallegni F., Tesi C., Ricci S. 2021. Bioarchaeology of the human remains of the so-called “sailor” from the site of the ancient ships of Pisa – San Rossore. Medicina Historica 3, 196-205.


Ricci S., Capecchi G., Boschin F., Arrighi S., Ronchitelli A., Condemi S., 2016, Toothpick use among Epigravettian humans from Grotta Paglicci (Italy). International Journal of Osteoarchaeology, 26(2): 281-289.


Benazzi S., Fornai C., Buti L., Toussaint M., Mallegni F., Ricci S., Gruppioni G., Weber G. W., Condemi S. and Ronchitelli A., 2012. Cervical and Crown Outline Analysis of Worn Neanderthal and Modern Human Lower Second Deciduous Molars. American Journal of Physical Anthropology 149/4, 537-546.


Condemi S., Tardivo D., Foti B., Ricci S., Giunti P., Longo L., 2012. A case of an osteolytic lesion on an Italian Neanderthal jaw.  C.R. Palevol 11, 79-83.


Fu Q., Mittnik A., Johnson P. L.F., Bos K., Lari M., Bollongino R., Sun C., Giemsch L., Schmitz R., Burger J., Ronchitelli A.M., Martini F., Cremonesi R.G., Svoboda J., Bauer P., Caramelli D., Castellano S., Reich D., Pääbo S. and Krause J., 2013. A Revised Timescale for Human Evolution Based on Ancient Mitochondrial Genomes. Current Biology 23, 553–559.

Accademia dei Fisiocritici - Mostra sul Riparo del Molare

For many years the RUPA has been collaborating with other institutions in the setting up of museum itineraries and exhibitions inspired by didactic and scientific aims (e.g: documentary/educational itinerary at the Quaternary Study Centre in Sansepolcro, Virtual Museum in Marina di Camerota, Prehistoric Section of the National Archaeological Museum in Perugia, Museum on Grotta Paglicci in Rignano Garganico, Civic Archaeological Museum in La Spezia, Civic Archaeological Museum in Camaiore, "The Colors of Darkness" exhibition set at the Vittoriano in Rome, "Darwin 1809 - 2009" exhibition in Bari, etc. ...).


Heritage offices of Toscana, Umbria, Lazio, Abruzzo, Campania, and Puglia.


Research Institutes

  • Berna F., Geoarchaeology (Simon Fraser University, British Columbia, Canada).
  • Parenti F., Prehistory (Universidade Federal do Paraná, Departamento de Antropologia - Curitiba - Brazil).
  • Higham T.F.G., Radiocarbon dating (Department of Evolutionary Anthropology, Faculty of Life Sciences, University of Vienna - Austria).
  • Douka K., Prehistory (Department of Evolutionary Anthropology, Faculty of Life Sciences University of Vienna - Austria).
  • Sano K., Use- wear traces and ballistic (Center for Northeast Asian Studies, Tohoku University Kawauchi 41, Aoba-ku, Sendai 980-8576, Japan).
  • Petrequin P., Neolithic and Calcolithic (French National Centre for Scientific Research – MSHE Franche-Comté).
  • Ruban D., Geology (Department of Business in Hospitality Sphere Higher School of Business Southern Federal University Rostov-na-Donu - Russian Federation).
  • Manica A., Evolutionary Ecology (Department of Zoology - University of Cambridge - UK).
  • Toskan B., Zooarchaeology (Istitute of Archaeology, ZRC-SAZU, Ljubljana – Slovenia).
  • Bahen J.J., Geochronology (Muséum National d'Histoire Naturelle, Paris, France).
  • Falgueres C., Geochronology (CNRS, UMR 7194, Histoire naturelle de l'Homme préhistorique, Paris, France).
  • Tomasso A., Prehistory (CNRS, UMR 7264, Cultures – Environnements. Préhistoire, Antiquité, Moyen Âge, Nice, France).
  • Saliari K., Zooarchaeology (Natural History Museum, Vienna, Austria).

Research Institutes

  • Benazzi S., Osteoarchaeology and Palaeoanthropology (Department of Cultural Heritage - University of Bologna - Italy).
  • Negrino F., Prehistory (Department of Antiquity, Philosophy  and History - University of Genova -Italy).
  • Benvenuti M., Geomorphology; Santo A.P. Archaeometry (Department of Earth Sciences –University of Florence - Italy).
  • Caramelli D.,Genetics (Department of Biology - University of Firenze - Italy).
  • Revedin A., Prehistory (Italian Institute of Prehistory and Protohistory).
  • Talamo S., Radiocarbon dating specialist (Department of Chemistry - University of Bologna - Italy).
  • Vaccari L., FTIR Microscopy and Spectroscopy  (Elettra Sincrotone Trieste - Italy).
  • Marini F., chemiometric methods for cultural heritage (Department of Chemistry - University of Rome "La Sapienza"- Italy).
  • De Rita F., Magri D., Paleobotany (Dipartimento di Biologia Ambientale, Sapienza, University of Roma).
  • Carra M., Paleobotany (Dipartimento di Storia, Culture, Civiltà, University of Bologna).
  • Bernardini F., Prehistory (Dipartimento di Studi Umanistici, Università Ca’ Foscari, Venezia).
  • Arbulla D., Prehistory (Museo Civico di Storia Naturale di Trieste).

Other Collaborations

  • Associazione degli Antropologi Italiani (AAI) - Gruppo di Biologia dello Scheletro.
  • Centro Studi sul Quaternario Onlus (CeSQ) di Sansepolcro (AR). Attività di orientamento, formazione, ricerca e studio (in convenzione con l'Università di Siena).
  • Istituto italiano di Preistoria e Protostoria (Firenze)
  • Istituto Italiano di Paleontologia Umana (Roma)
  • Municipalities of Sansepolcro (AR), Monte Argentario and Roccastrada(GR), Camerota and Castelcivita (SA), Rignano Garganico (FG), Bisceglie (BT), and Ginosa (TA).
  • Comando Provinciale dei Vigili del Fuoco di Grosseto e Ufficio Circondariale Marittimo di Porto Santo Stefano.


Stefano Ricci (Technician)

Vincenzo Spagnolo (Research Fellow)

Jacopo Crezzini (Research Fellow)

Clarissa Dominici (PhD Student)

Matteo Rossini (PhD Student)

Annamaria Ronchitelli (visiting fellow)

Paolo Boscato (visiting fellow)

Sem Scaramucci (visiting fellow)